Ingie Hong^1,2,25*, Juhyun Kim^1,3,4,25, Thomas Hainmueller^5,21,25, Dong Won Kim^1,22,23, Joram Keijser^6,7, Richard C. Johnson^1,2, Soo Hyun Park^8,9, Nathachit Limjunyawong^1,24, Zhuonan Yang^1,2, David Cheon^1,2, Taeyoung Hwang^1,10,11, Amit Agarwal^1,12,13, Thibault Cholvin⁵, Fenna M. Krienen¹⁴, Steven A. McCarroll¹⁵, Xinzhong Dong^1,16, David A. Leopold8, Seth Blackshaw^1,2,17,18, Henning Sprekeler^6,19,20, Dwight E. Bergles^1,2, Marlene Bartos⁵, Solange P. Brown^1,2 & Richard L. Huganir^1,2*

¹The Solomon H. Snyder Department of Neuroscience, Johns Hopkins University School of Medicine, Baltimore, MD, USA. 

²Kavli Neuroscience Discovery Institute, Johns Hopkins University, Baltimore, MD, USA. 

³Department of Psychiatry and Behavioral Sciences, Johns Hopkins University School of Medicine, Baltimore, MD, USA. 

⁴Emotion, Cognition and Behavior Research Group, Korea Brain Research Institute (KBRI), Daegu, Republic of Korea. 

⁵Institute for Physiology I, University of Freiburg, Medical Faculty, Freiburg, Germany. 

⁶Modelling of Cognitive Processes, Technical University of Berlin, Berlin, Germany. 

⁷Charité–Universitätsmedizin Berlin, Einstein Center for Neurosciences Berlin, Berlin, Germany. 

⁸Section on Cognitive Neurophysiology and Imaging, National Institute of Mental Health, Bethesda, MD, USA. 

⁹Department of Brain and Cognitive Sciences, Korea Advanced Institute of Science and Technology (KAIST), Daejeon, Republic of Korea. 

¹⁰Lieber Institute for Brain Development, Johns Hopkins Medical Campus, Baltimore, MD, USA. 

¹¹Department of Neurology, Johns Hopkins University School of Medicine, Baltimore, MD, USA. 

¹²Chica and Heinz Schaller Research Group, Institute for Anatomy and Cell Biology, Heidelberg, Germany. 

¹³Interdisciplinary Center for Neurosciences, University of Heidelberg, Heidelberg, Germany. 

¹⁴Princeton Neuroscience Institute, Princeton University, Princeton, NJ, USA. 

¹⁵Department of Genetics, Harvard Medical School, Boston, MA, USA. 

¹⁶Howard Hughes Medical Institute, Johns Hopkins University School of Medicine, Baltimore, MD, USA. 

¹⁷Department of Ophthalmology, Johns Hopkins University School of Medicine, Baltimore, MD, USA. 

¹⁸Institute for Cell Engineering, Johns Hopkins University School of Medicine, Baltimore, MD, USA. 

¹⁹Bernstein Center for Computational Neuroscience Berlin, Berlin, Germany. 

²⁰Science of Intelligence, Research Cluster of Excellence, Berlin, Germany. 

²¹Present address: Department of Psychiatry, New York University Langone Medical Center, New York, NY, USA. 

²²Present address: Danish Research Institute of Translational Neuroscience (DANDRITE), Nordic EMBL Partnership for Molecular Medicine, Aarhus University, Aarhus, Denmark. 

²³Present address: Department of Biomedicine, Aarhus University, Aarhus, Denmark. 

²⁴Present address: Center of Research Excellence in Allergy and Immunology, Research Department, Faculty of Medicine Siriraj Hospital, Mahidol University, Bangkok, Thailand. 

²⁵These authors contributed equally: Ingie Hong, Juhyun Kim, Thomas Hainmueller. 

*Corresponding authors

Correspondence to Ingie Hong or Richard L. Huganir.