Don-Kyu Kim ^{‡ 1}, Dongryeol Ryu ^{§ 1}, Minseob Koh ^{¶ 2}, Min-Woo Lee ^§, Donghyun Lim ^‖, Min-Jung Kim ^§, Yong-Hoon Kim ^**, Won-Jea Cho ^‡‡, Chul-Ho Lee ^**, Seung Bum Park ^{¶ ‖}, Seung-Hoi Koo ^§, Hueng-Sik Choi ^{‡ §§}

^‡National Creative Research Initiatives Center for Nuclear Receptor Signals, Hormone Research Center, School of Biological Sciences and Technology, Chonnam National University, Gwangju 500-757, Republic of Korea
^‡‡College of Pharmacy and Research Institute of Drug Development, Chonnam National University, Gwangju 500-757, Republic of Korea
^§Department of Molecular Cell Biology and Samsung Biomedical Research Institute, Sungkyunkwan University School of Medicine, 300 Chunchun-dong, Jangan-gu, Suwon, Gyeonggi-do 440-746, Republic of Korea
^¶Departments of Chemistry, Seoul National University, Seoul 151-747, Republic of Korea
^‖Departments of Biophysics and Chemical Biology, College of Natural Sciences, Seoul National University, Seoul 151-747, Republic of Korea
^**Korea Research Institute of Bioscience and Biotechnology, Daejeon 305-806, Republic of Korea
^§§Research Institute of Medical Sciences, Department of Biomedical Sciences, Chonnam National University Medical School, Gwangju 501-746, Republic of Korea

¹Both authors contributed equally to this work.
²Supported by a Brain Korea 21 Program fellowship award and a Seoul Science fellowship award.
^*Corresponding authors: Seung Bum Park, Seung-Hoi Koo or Hueng-Sik Choi